When uncompressed, the file is still 4.3gb because it contains loads of 'filler' data, again this is garbage but now it's a type of garbage that is archive friendly. Then it overwrites the garbage areas with a uniform value (0xFF). Just like the Wii, it reads the partition table to determine where the game area is, and which areas are garbage (unused). The reason is, all that garbage has to be compressed as well, which is basically wasted space, time, and bandwidth. However due to this garbage being there, when you try to compress the ISO you'll get a marginally smaller file (perhaps 4.1gb instead of 4.3gb).
The garbage areas are ignored by the Wii, since the partition table doesn't have any entries for those areas so the Wii knows to stay away. The partition table on the disc tells the Wii where to look on the disc for actual game data. The unused portion of the disc is filled with garbage. Some games only use 10% of the total DVD capacity, it ranges basically from 10% to 100%. For some games (such as AC:CF and many others), only a little bit of actual disc space is required for the game files.
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